Did you also pull the ROM SIMM that has the postscript interpreter on it? I don't think the JetDirect card will do much good without it, unless HP released PCL5 drivers for MacOS. (Never heard of any.)
http://www.ebay.com/itm/C3129A-C3129-60001-LaserJet-4-4Plus-PostScript-Simm-Kit-w-16MB-MEMORY-UPGRADE-/281729237990?hash=item41985f03e6